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The year is 2028. The Rebellion Army, under the control of General Morden, is seeking world domination through military might and has captured much of the Regular Army's soldiers and equipment, including a newly designed tank, code named "Metal Slug." Marco Rossi and Tarma Roving (with other characters appearing in later titles) must fight to stop General Morden's ambitions, finding help from whatever soldiers they can rescue and any equipment they can commandeer. If it can't be taken, destroy it!

...But let's get serious now - plots in most action games just don't matter. Who really remembers the whys and wherefores of Contra? Does it really matter what Dr. Wiley is up to in the NES Mega Man games? Likewise, the background story to the Metal Slug series doesn't really matter. It's just you against the world and your reflexes are your only friend.